PCIe x1 x4 cards in x16 slot

can you put a pcie x4 in a x16 slot
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I did contact the card manufacturer, and they said that putting a x4 card in x16 slot is called up-pluging, and they know for sure that these

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wild gems slot A GTX 780R9 290 for example may suffer a performance hit but an R7 260GTX 750 may not The PCIe slots are rated as they are because of I explain in this video the Types of PCIe slots, how you can identify them, and also what their